North American Recent Soft-shelled Turtles (Family  Trionychidae)

University of Kansas Publications, Museum of Natural History 13 (10): 429-611.

Summary (Auszug):

In North America, soft-shelled turtles (genus Trionyx) occur in northern México, the eastern two-thirds of the United States, and extreme southeastern Canada. The genus fits the well-known Sino-American distributional pattern. In North America there are four species. Three (ferox, spinifer and muticus) are well-differentiated [591] and one (ater) is not well-differentiated from spinifer. Characters of taxonomic worth are provided by the following: size; proportions of snout, head and shell; pattern on carapace, snout, side of head, and limbs; tuberculation; sizes of parts of skull; number of parts of carapaces; and, shape and number of some parts of plastra. Many features show geographical gradients or clines. T. ferox is the largest species and muticus is the smallest. Females of all species are larger than males. With increasing size of individual, the juvenal pattern is replaced by a mottled and blotched pattern in females of all species; adult males of spinifer retain a conspicuous juvenal pattern, whereas the juvenal pattern is sometimes obscured or lost on those of ferox and muticus. The elongation of the preanal region in all males, and the acquisition of a "sandpapery" carapace in males of spinifer occur at sexual maturity. There is a marked secondary sexual difference in coloration in a population of T. s. emoryi (side of head bright orange in males and yellow in females). The sex of many hatchlings of T. s. asper can be distinguished by the pattern on the carapace. Slight ontogenetic variation occurs in some proportional measurements. Large skulls of ferox and some asper (those in Atlantic Coast drainages) have expanded crushing surfaces on the jaws. Considering osteological characters, muticus is most distinct; there is less difference between ferox and spinifer than between those species and muticus.